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In a stable steady state, " $ \kappa \ne \kappa'$"

We now show that there is no stable steady state with $ r_n=\max_i r_i$. Suppose that $ Q_n>0$, $ Q_{n-1}<0$, and $ r_n=\max_i r_i$. Then

$\displaystyle P(0)=\Pi_{j\ne n}Q_j/P_j + Q_n/P_n \Sigma_{i=1}^{n-1} \Pi_{j=1,j\ne i}^{n-1}Q_j/P_j -\Pi_{j=1}^{n}Q_j/P_j
$

$\displaystyle P(0)=\left( \Pi_{j\ne n}Q_j/P_j + Q_n/P_n \Pi_{j=1}^{n-2} Q_j/P_j...
...Q_n/P_n \Sigma_{i=1}^{n-2} \Pi_{j=1,j\ne i}^{n-1}Q_j/P_j -\Pi_{j=1}^{n}Q_j/P_j
$

$\displaystyle P(0)=\left( Q_{n-1}/P_{n-1} + Q_n/P_n\right) \Pi_{j=1}^{n-2}Q_j/P...
...Q_n/P_n \Sigma_{i=1}^{n-2} \Pi_{j=1,j\ne i}^{n-1}Q_j/P_j -\Pi_{j=1}^{n}Q_j/P_j
$

The last two terms in the sum both have the same sign as $ (-1)^n$.

Now consider

$\displaystyle \frac{Q_n}{P_n}\frac{P_{n-1}}{Q_{n-1}} = \frac{1-2\beta_n x_n}{1-2\beta_{n-1} x_{n-1}} \frac{\beta_{n-1}}{\beta_n}
$

$\displaystyle \frac{Q_n}{P_n}\frac{P_{n-1}}{Q_{n-1}} = - \frac{\sqrt{1-4 \alpha...
... \beta_n^2}} {\sqrt{1-4 \alpha D^2 \beta_{n-1}^2}} \frac{\beta_{n-1}}{\beta_n}
$

By hypothesis, $ \frac{\beta_{n-1}}{\beta_n}\ge1$, and thus

$\displaystyle Q_n/P_n\ge \left\vert {Q_{n-1}/P_{n-1}} \right\vert
$

Therefore, $ P(0)$ has the same sign as $ (-1)^n$. Since $ P(Q_n)$ has the same sign as $ (-1)^{n-1}$, P has a positive root, and the steady state is unstable.



Cinquin & Page, Bull Math Biol (2006, in press)